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29

Ungulates have some of the longest migrations of all terrestrial

animals, up to several thousand kilometres for species such as

the North American caribou (

Rangifer

ssp.). Migration is a cru-

cial element in the survival of many ungulates, their ability to

survive in marginal landscapes being based on the opportunity

to migrate. Twenty-four large mammal species (and their sub-

species) are known to migrate in large aggregations today – all

of these are ungulates (Harris

et al.

, 2009).

Infrastructure may have an impact on wild ungulates by creat-

ing direct disturbance and road kills locally, though this effect

is usually less important compared with avoidance or blocking

of migrations. Of far greater concern is when infrastructure

generates increased traffic and human activity surrounding

these corridors leading to increased logging, hunting, poach-

ing and settlements, as well as introduction of invasive species,

livestock and agricultural expansion. This in turn, may lead to

more regional indirect impacts such as avoidance of road cor-

ridors in the range of 4–10 km, and even up to 30 km, by mi-

grating ungulates, thus generating semi-permeable corridors.

These are corridors that in theory are passable, but rarely, de-

pending on the situation at hand, are crossed in reality. The

combined actions lead to cumulative impacts, resulting in a

partial or full breakdown of the ecological network involved,

Road development and agricultural expansion