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23

Tundra

Boreal forest

Temperate forest

Temperate grasslands, savannas and shrublands

Desert and dry shrublands

Tropical and subtropical forests

Tropical and subtropical grasslands, savannas and shrublands

Source: adapted from Olson

et al

., 2001.

Much of the original area of temperate grassland has been cleared for agriculture.

Where natural vegetation remains, minimising human disturbance can prevent fur-

ther carbon loss.

Grasslands are found across much of the world as an early succes-

sional ecosystem in forested regions. They also form the natural

vegetation in climates where precipitation levels are inadequate

to support trees but higher than those of deserts (Woodward

et

al.

2004). Extensive areas of natural temperate grassland occur in

South America, the USA and Central Asia. Plant growth in these

grasslands is water and nutrient limited and plants allocate much

of their biomass below ground, where they produce slowly decom-

posing roots. Grazing animals typically play an important role in

maintaining grasslands in that they accelerate carbon cycling by

consuming and respiring large quantities of leaf biomass and re-

turning some of this to the soil as dung. This is a form of organic

carbon that is more decomposable than the leaf and root litter of

grasses. In many areas this role is now performed by livestock.

Overall, temperate grasslands have low levels of plant biomass

compared with forest or shrubland ecosystems (e.g. 0.68 and

7.3 t C per ha respectively in the temperate steppe of China,

Fan

et al.

2008). However, their soil organic carbon stocks tend

to be higher than those of temperate forests (133 t C per ha,

Amundson 2001).

HUMAN IMPACTS AND IMPLICATIONS FOR

CARBON MANAGEMENT

Despite only having intermediate productivity some temperate

grasslands are well suited to crop production and can produce

excellent agricultural soils. In much of their natural range, e.g.

the prairies of America, these have been cleared to make way

for intensive agriculture.

TEMPERATE GRASSLANDS