246 / 1173
0.2
0.3
0.4
0.5
0.6
0.7
0.8
0.9
1
0
1
2
3
4
5
6
Dose/Gy
Surviving fraction
α
s
α
r
S
=
e
−
α
D
−
β
D
2
α
=
α
r
1
+
α
s
α
r
−
1
(
)
e
−
D D
c
(
)
D
c
Low-dose
hyper-
radiosensitivity
First reported in 1986 in
mouse epidermis and kidney
Short S, Mayes C, Woodcock M,
Johns H, Joiner MC (1999).
Int J Radiat Biol 75: 847–55.
T98G human
GBM cells